This sparkling, genre-bending novel opens with amateur historian Anne, who has a passion for research into the murkier corners of England in the 1600s. In an archive, Anne has stumbled across an obscure memoir, one that hints at an intricate tapestry of secret lives and loves.

The full story eventually weaves together five manuscripts, each a different thread in the same strange tale: The Plague descends upon a village, and two children, Joan and Thomasina, are the only survivors. They bond with each other and with "Old Nut," a woman who lives in the forest nearby. But when relatives return, Old Nut is accused of witchcraft and condemned to death. Joan is hired as a maid to well-educated Lady Margaret Long—and, being lively and curious, soon becomes a beloved companion. Thomasina is sent on a perilous voyage to Virginia, where she adopts boys' clothing and navigates life as a male.

Years later, Tom and Joan find each other and fall in love—but are discovered, naked, by a clergyman. Horrified, he believes there can only be one explanation for Tom's "unmanned" state: Joan is a witch and, like Old Nut years ago, must be tried for sorcery. It falls upon Anne, reading between faded pages and centuries, to uncover the fate of the lovers—and add her own contemporary line of "truth" to this tale from a time when there were no labels for who Tom and Joan might be.